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Struthiopteris spicant

No recorded common name

The hard-fern grows two strikingly different kinds of fronds on a single plant, a trait that sets it apart from many other ferns in its family, the Blechnaceae.

Also known by the common names hard-fern and deer fern, it belongs to the order Polypodiales within the class Polypodiopsida. It is evergreen and widespread across temperate regions of the Northern Hemisphere, making it a familiar sight in woodland and mountain settings where conditions stay cool and moist year-round.

Identification and Appearance

The clearest way to recognize Struthiopteris spicant is by its two distinct frond types growing from the same plant. Sterile fronds lie flatter and spread outward, with flat, wavy-margined leaflets measuring 5 to 8 millimeters wide. Fertile fronds stand more upright and bear much narrower leaflets, each marked underneath by two thick rows of sori, the spore-producing structures that distinguish them at a glance from the sterile growth.

This dimorphism, the pairing of two leaf forms with different jobs, is the fern's most distinctive feature and makes it relatively easy to separate from similar-looking ferns that produce only one uniform type of frond.

Distribution and Habitat

Struthiopteris spicant has a wide native range spanning Macaronesia, northwestern Africa, Europe, and east to Iran, with a separate presence from Alaska south through western Canada into California, Idaho, Oregon, and Washington. Within Europe it is recorded from Iceland and Scandinavia south to the Mediterranean islands of Corsica, Sardinia, Sicily, and Crete, and east through the Baltic states, Ukraine, and the Caucasus.

The species favors cool, moist, temperate conditions and is hardy down to -20°C. Its evergreen habit lets it persist through winter in these climates, reaching a height of about 0.5 meters.

Observation records through the year show a pronounced rise from spring into a peak around midsummer, tapering off toward late autumn and winter, consistent with a plant that is most visible and actively growing during the warmer months, though such records reflect observation activity rather than the fern's actual abundance or full range.

World map showing recorded occurrences of Struthiopteris spicant.
Where it has been recorded · GBIF
44,498
JanFebMarAprMayJunJulAugSepOctNovDec
Records by month, all years · GBIF

Growth and Form

Struthiopteris spicant is a perennial fern that reaches roughly 0.5 meters in height. Its evergreen foliage persists through the year, allowing the plant to remain visible and green even through cold winters in its hardy range down to -20°C.

Uses and Human Relationships

The root, shoots, and leaves of Struthiopteris spicant have been used as a famine food, cooked for consumption in times of scarcity. Beyond subsistence use, the plant has ornamental value: it has received the Royal Horticultural Society's Award of Garden Merit, a recognition of its qualities as a garden plant.

Fun Facts

  1. The species was first described in 1753 by Carl Linnaeus, who classified it as Osmunda spicant.
  2. Since then it has been shuffled through a wide range of genera, including Blechnum, before being placed in Struthiopteris under the Pteridophyte Phylogeny Group classification of 2016 (PPG I).

Photos

Photographs from by Emerson via iNaturalist, licensed under CC BY 4.0., via iNaturalist, licensed under CC0 1.0., by John Brew via iNaturalist, licensed under CC BY 4.0., by giantcicada via iNaturalist, licensed under CC BY 4.0., by Timothy McNitt via iNaturalist, licensed under CC BY 4.0., by Andrejus Gaidamavičius via iNaturalist, licensed under CC BY 4.0., by Alexis Tinker-Tsavalas via iNaturalist, licensed under CC BY 4.0., by Adam Górski via iNaturalist, licensed under CC BY 4.0., by Gabriele Vaudano via iNaturalist, licensed under CC BY 4.0., by Cass Reaume via iNaturalist, licensed under CC BY 4.0., and by Scott Shanks via iNaturalist, licensed under CC BY 4.0., credited on each photo.

Ecology and classification

Growth habit

  • Fern

Classification

Phylum
Tracheophyta
Class
Polypodiopsida
Order
Polypodiales
Family
Blechnaceae
Genus
Struthiopteris
Species
Struthiopteris spicant

Sources

  1. GBIF Species names and classification
  2. iNaturalist Photos
  3. Wikidata Common names and conservation status
  4. Wikipedia Summary extract
  5. Plants of the World Online Native range, life form and climate (WCVP)
  6. GloBI What it eats, what eats it, parasites and pollinators
  7. MammalBase